Ademosynidae

Ademosynidae is an extinct family of beetles, known from the Late Triassic to Early Cretaceous. There are at least six genera and 11 species in Ademosynidae. Members of the family were small oval beetles, with a length generally smaller than . Characteristics of the family include a pronotum without anterior angles and a rounded anterior margin, and elytra with 9–12 punctate striae. The systematic position of the family within the order Coleoptera is currently uncertain: they can be considered early examples of the suborder Polyphaga, or unusual examples of Archostemata. Ademosynidae has sometimes been synonymized with the older family Permosynidae, even though the latter was established based on fossils of isolated elytra and the former was established based on fossils of complete bodies.

Genera

The family Ademosynidae is restricted to the following six genera and 11 species in Yan, Beutel and Ponomarenko (2017):

• † Ademosyne <small>Handlirsch, 1906</small>
• †Ademosyne bacca <small>Ponomarenko, 1969</small> – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
• †Ademosyne elliptica <small>Ponomarenko, 1969</small> – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
• †Ademosyne kirghizica <small>Ponomarenko, 1969</small> – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
• †Ademosyne major <small>Handlirsch, 1906</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• † Cephalosyne <small>Ponomarenko, 1969</small> – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
• †Cephalosyne capitata <small>Ponomarenko, 1969</small>
• † Dolichosyne <small>Ponomarenko, 1969</small> – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
• †Dolichosyne confragosa <small>Ponomarenko, 1969</small>
• †Dolichosyne rostrata <small>Ponomarenko, 1969</small>
• †Dolichosyne sulcata <small>Ponomarenko, 1969</small>
• † Gnathosyne <small>Ponomarenko, 1969</small> – Tologoi Formation, Kazakhstan, Late Triassic (Norian)
• †Gnathosyne akkolkensis <small>Ponomarenko, 1969</small>
• † Petrosyne <small>Ponomarenko, 1969</small> – Kyrgyzstan, Early Jurassic (Pliensbachian)
• †Petrosyne liassica <small>Ponomarenko, 1969</small>
• † Sphaerosyne <small>Ponomarenko, 1969</small> – Zaza Formation, Russia, Early Cretaceous (Aptian)
• †Sphaerosyne globosa <small>Ponomarenko, 1969</small>

The following genera and species are also included in the family according to Fossilworks, but are excluded or not mentioned in Yan, Beutel and Ponomarenko (2017). Some of these species, for instance those of Ademosynoides and those of Ademosyne described by Tillyard and Dunstan, were described from fossils of isolated elytra.

• † Ademosynoides <small>Dunstan, 1923</small>
• †Ademosynoides abnormis <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides alternata <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides angusta <small>(Tillyard, 1916)</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides antarctica <small>Zeuner, 1959</small> – Mount Flora Formation, Antarctica, Middle Jurassic (Bathonian)
• †Ademosynoides japonicus <small>Fujiyama, 1973</small> – Mononoki Formation, Japan, Late Triassic (Carnian)
• †Ademosynoides magnifica <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides minor <small>(Handlirsch, 1906)</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides obtusa <small>(Tillyard, 1916)</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †Ademosynoides striatella <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• † Atalosyne <small>Ren, 1995</small> – Lushangfen Formation, China, Early Cretaceous (Aptian)
• †Atalosyne sinuolata <small>Ren, 1995</small>
• † Chaocoleus <small>Ponomarenko, Yan & Huang, 2014</small> – Yinping Formation, China, Permian (Capitanian)
• †Chaocoleus limnebius <small>Ponomarenko, Yan & Huang, 2014</small>
• † Eremisyne <small>Wang, 1998</small> – Xiazhuang Formation, China, Early Cretaceous (Aptian/Albian)
• †Eremisyne xiazhuangensis <small>Wang, 1998</small>
• † Grammositus <small>Dunstan, 1923</small> (also spelled Grammositum) – Blackstone Formation, Australia, Late Triassic (Norian)
• †Grammositus bilineatus <small>Dunstan, 1923</small>
• †"Ademosyne" adunca <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" arcucciae <small>Martins-Neto and Gallego, 2006</small> – Los Rastros Formation, Argentina, Late Triassic (Carnian)
• †"Ademosyne" australiensis <small>Tillyard, 1916</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" brevis <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" cameroni <small>Tillyard, 1916</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" congener <small>Tillyard, 1916</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" curvata <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" elongatus <small>Martins-Neto and Gallego, 2006</small> – Los Rastros Formation, Argentina, Late Triassic (Carnian)
• †"Ademosyne" hexacostata <small>Martins-Neto and Gallego, 2006</small> – Los Rastros Formation, Argentina, Late Triassic (Carnian)
• †"Ademosyne" intermedia <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" lata <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" llantenesensis <small>Brauckmann et al., 2010</small> – Llantenes Formation, Argentina, Late Triassic (Norian)
• †"Ademosyne" olliffi <small>(Handlirsch, 1906)</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" parva <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" polyzetete <small>Ponomarenko, 2011</small> – Lunz Formation, Austria, Late Triassic (Carnian)
• †"Ademosyne" prisca <small>Riek, 1974</small> – Molteno Formation, South Africa, Late Triassic (Carnian)
• †"Ademosyne" punctata <small>Tillyard, 1916</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" punctuada <small>Martins-Neto and Gallego, 2006</small> – Los Rastros Formation, Argentina, Late Triassic (Carnian)
• †"Ademosyne" ramocostata <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" reducta <small>Riek, 1976</small> – Molteno Formation, South Africa, Late Triassic (Carnian)
• †"Ademosyne" rosenfeldi <small>Brauckmann et al., 2010</small> – Llantenes Formation, Argentina, Late Triassic (Norian)
• †"Ademosyne" rugulosa <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" sibirica <small>Rohdendorf, 1961</small> – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian (Wuchiapingian/Changhsingian)
• †"Ademosyne" speciosa <small>Riek, 1974</small> – Molteno Formation, South Africa, Late Triassic (Carnian)
• †"Ademosyne" umutu <small>Martins-Neto and Gallego, 2009</small> – Los Rastros Formation, Argentina, Late Triassic (Carnian)
• †"Ademosyne" vittamargina <small>Dunstan, 1923</small> – Blackstone Formation, Australia, Late Triassic (Norian)
• †"Ademosyne" wianamattensis <small>Tillyard, 1918</small> – Ashfield Shale, Australia, Middle Triassic (Pelsonian/Illyrian)

References